| 1 | | Harvard University |
| 2 | | Stanford University |
| 3 | | Massachusetts Institute of Technology |
| 4 | | University of California, Berkeley |
| 5 | | University of Cambridge |
| 6 | | Princeton University |
| 7 | | California Institute of Technology |
| 8 | | Columbia University |
| 9 | | University of Oxford |
| 9 | | University of Chicago |
| 11 | | Yale University |
| 12 | | University of California, Los Angeles |
| 13 | | Cornell University |
| 14 | | University of California San Diego |
| 15 | | University of Washington |
| 16 | | University of Pennsylvania |
| 17 | | Johns Hopkins University |
| 18 | | University of California, San Francisco |
| 19 | | ETH Zurich |
| 20 | | University College London |
| 21 | | The University of Tokyo |
| 22 | | University of Michigan |
| 22 | | Imperial College London |
| 24 | | University of Wisconsin–Madison |
| 24 | | University of Toronto |
| 26 | | Kyoto University |
| 27 | | New York University |
| 28 | | Northwestern University |
| 28 | | Lemann Center for Brazilian Studies |
| 30 | | University of Minnesota |
| 31 | | Duke University |
| 32 | | Washington University in St. Louis |
| 33 | | Rockefeller University |
| 34 | | University of Colorado Boulder |
| 35 | | Biologie Computationnelle, Quantitative et Synthétique |
| 36 | | University of North Carolina at Chapel Hill |
| 37 | | University of British Columbia |
| 38 | | University of Manchester |
| 39 | | University of Copenhagen |
| 39 | | The University of Texas at Austin |
| 41 | | University of California, Santa Barbara |
| 42 | | Université Paris-Sud |
| 43 | | University of Maryland, College Park |
| 44 | | The University of Melbourne |
| 45 | | The University of Texas Southwestern Medical Center |
| 45 | | University of Edinburgh |
| 47 | | University of California, Irvine |
| 47 | | Karolinska Institutet |
| 49 | | Heidelberg University |
| 49 | | Ludwig-Maximilians-Universität München |
